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5 Conclusion Through integrative transcriptomic-proteomic analysis, this study revealed that winter nutritional supplementation remodels yak cardiac molecular regulatory networks by activating selenoprotein-mediated antioxidant systems (significant upregulation of GPX1 and SELENOW ) and downregulating immune surveillance molecules ( ULBP family and GZMB ), with post-transcriptional regulation as the predominant mechanism (87.3% protein-specific changes)

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